Emma Darwin, widow of scientist Charles Darwin, finds herself on a late life journey of self-discovery as she faces a challenge to her faith in the form of her beloved atheist husband’s as–yet unpublished autobiography.
| Actor | Role |
|---|---|
 Anthony HopkinsCharles Darwin | Charles Darwin |
 Charlotte RamplingEmma Darwin (née Wedgwood) | Emma Darwin (née Wedgwood) |
 Tom HollanderMarshall Winwick | Marshall Winwick |
 Billy HowleGeorge Darwin | George Darwin |
 Mia McKenna-BruceJane | Jane |