A psychological, heart-wrenching love story that provides a unique and inside look at Charles Darwin. Torn between faith and science, he struggles to finish his legendary book "On the Origin of Species," which goes on to become the foundation for evolutionary biology.
| Actor | Role |
|---|---|
 Paul BettanyCharles Darwin | Charles Darwin |
 Jennifer ConnellyEmma Darwin | Emma Darwin |
 Martha WestAnne Darwin | Anne Darwin |
 Guy HenryTechnician | Technician |
 Jeremy NorthamJohn Brodie-Innes | John Brodie-Innes |
 Toby JonesThomas Henry Huxley | Thomas Henry Huxley |
 Benedict CumberbatchJoseph Dalton Hooker | Joseph Dalton Hooker |
 Jim CarterParslow | Parslow |
 Robert GlenisterDr. Holland | Dr. Holland |
 Ian MercerGoodman | Goodman |
 Bill PatersonDr. Gully | Dr. Gully |
| TTeresa ChurcherMrs. Davies | Mrs. Davies |
 Richard RidingsThatcher | Thatcher |
 Freya ParksEtty Darwin | Etty Darwin |
| IIan KellyCaptain Fitzroy | Captain Fitzroy |
| AAnabolena RodriguezFuegia Basket | Fuegia Basket |
| PPaul CampbellBoat Memory | Boat Memory |
| ZZak DaviesJemmy Button | Jemmy Button |
| CChristopher DunkinGeorge Darwin | George Darwin |
 Gene GoodmanFranky Darwin | Franky Darwin |
| HHarrison SansostriLenny Darwin | Lenny Darwin |